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Comment: Revised based on observations in the UAT tier.

Specify If you clicked the download icon in the row for one collection, you can specify a destination hierarchy:

OptionDescription
Download to Specified FolderDownloads to the destination point all the files from the specified collection and its subfolders.
Create Parent FolderDownloads as stated above and also copies the parent folder. 
Create Full PathDownloads as stated above and also copies the full path.